M

macroalgae The multicellular brown, red, and green algae.

macroelements Essential elements needed for plant growth in relatively large amounts; C, H, O, P, K, N, S, Ca, Fe, Mg.

macroevolution Evolutionary change on a grand scale, encompassing the origin of novel designs, evolutionary trends, adaptive radiation, and mass extinction.

macrofauna benthic animals larger than about 0.5 mm

macrofibril An aggregation of microfibrils in the cell wall, visible with the light microscope.

macrogamete The larger of the two gamete types in a heterogametic organism, considered the female gamete.

macromere The largest size class of blastomeres in a cleaving embryo when the blastomeres differ in size from one another.

macromolecule A very large molecule, generally used in reference to carbohydrates, lipids, proteins, and nucleic acids.

macronucleus The larger of the two kinds of nuclei in ciliate protozoa; controls all cell functions except reproduction.

macronutrient An essential nutrient for which an animal has a large minimal daily requirement (greater than 100 mg) (e.g., calcium, phosphorus, magnesium, potassium, sodium, and chloride along with carbohydrates, lipids, and proteins).

macroparasite Large parasite that does not multiply in the host of interest. Examples are cestodes, trematodes, and most nematodes in their definitive hosts.

macrophage A phagocytic cell type in vertebrates that performs crucial functions in the immune response and inflammation, such as presenting antigenic epitopes to T cells and producing several cytokines.

macrophage migration inhibitory factor (MIF) Cytokine released by sensitized lymphocytes that tends to inhibit migration of macrophages in the immediate vicinity, thus contributing to accumulation of larger numbers of macrophages close to the site of MIF release.

macrophytes See seaweeds.

macroplankton The component of the plankton that consists of large organisms 2 to 20 cm in size.

macropores Large pore spaces caused by invertebrates and larger animals, including reptiles and mammals, that permeate the soil. They drain water not held by capillarity.

madreporite Sievelike structure that is the intake for the water-vascular system of echinoderms.

magma Molten rock from deep in the earth's interior; called lava when it spews from volcanic vents.

magnetic anomalies Magnetic bands in the sea floor that run parallel to the mid-ocean ridge .

magnetic confinement A technique for enclosing a nuclear fusion reaction in a powerful magnetic field inside a vacuum chamber.

magnetite A magnetic, iron-containing material.

main thermocline See thermocline.

major histocompatibility complex (MHC). Complex of genes coding for proteins inserted in the cell membrane; the proteins are the basis of self-nonself recognition by the immune system.

major mineral Mineral requirement needed in relatively large amounts.

Malacostraca The class of crustaceans whose members are characterized by having appendages modified for crawling along the substrate, as in lobsters, crayfish, and crabs. Alternatively, the abdomen and body appendages may be used in swimming, as in shrimp.

malacostracan Any member of the crustacean subclass Malacostraca, which includes both aquatic and terrestrial forms of crabs, lobsters, shrimps, pillbugs, sand fleas, and others.

malar mala = the jaw, cheek.

malate Four-carbon organic acid found with aspartate to be the first products of the C4 pathway of photosynthesis.

male dominance polygyny A mating system in which males compete and acquire dominance ranks that influence their access to females, with higher-ranking males often having greater mating activity.

male parasitism The permanent attachment of a male to a female in some deep-sea fishes.

malignant tumor A mass of cancerous cells that have left their site of origin, migrated through the body, invaded normal tissues, and are growing out of control.

malleus The ossicle attached to the tympanum in middle ears of mammals.

malnourishment A nutritional imbalance caused by lack of specific dietary components or inability to absorb or utilize essential nutrients.

malpighian tubules (Marcello Malpighi, Italian anatomist, 1628-1694). Blind tubules opening into the hindgut of nearly all insects and some myriapods and arachnids, and functioning primarily as excretory organs.

Malthusian growth A population explosion followed by a population crash; also called irruptive growth.

mamelon Ventral, serrated projection on the ventral surface of a male nematode of the family Syphaciidae. Its function is unknown.

Mammalia The class of vertebrates whose members are at least partially covered by hair, have specialized teeth, and are endothermic. Young are nursed from mammary glands. The mammals.

Mammalian Pertaining to animals feeding their young with milk from a breast.

mammals Vertebrates that have hair and mammary glands.

mammary gland The breast. In female mammals, the mammary glands produce and secrete milk to nourish developing young.

Man and Biosphere (MAB) program A design for nature preserves that divides protected areas into zones with different purposes. Critical ecosystem functions and endangered wildlife are protected in a central core region where limited scientific study is the only human access allowed. Ecotourism and research facilities are located in a relatively pristine buffer zone around the core, while sustainable resource harvesting and permanent habitation are allowed in multiple-use peripheral regions.

mandibles Third pair of appendages from the anterior in Crustacea; second pair in Insecta; primarily function in feeding; derived from appendages on primitive fourth (first postoral) somite.

mangal a tropical community of mangrove plants and associated organisms

manganese nodules Lumps of minerals (including manganese and other valuable minerals) that are found on the sea floor beyond the continental shelf.

mange Dermatitis caused by species of mites, often designated with the causative organism. For example, Sarcoptes causes sarcoptic mange.

mangroves Shrubs and trees that live along the seashore in tropical and subtropical regions and tolerate inundation by seawater.

mantle cavity The space between the mantle and the visceral mass of molluscs.

mantle (1) The semi-liquid region between the crust and core of the earth. (2) The outer layer of tissue that secretes the shell of molluscs.

manubrium The portion projecting from the oral side of a jellyfish medusa, bearing the mouth; oral cone; presternum or anterior part of sternum; handle-like part of malleus of ear.

marasmus Malnutrition, especially of infants, caused by a diet deficient in both calories and protein.

margin The edge of a flattened structure; in leaves, the lateral edge of the blade.

marginal bodies Sensory pits or short tentacles between the marginal loculi of the opisthaptor of an aspidogastrean trematode.

marginal meristem The meristem along the margin of a leaf primordium responsible with the formation and shape of the leaf.

marginal value theorem A model that predicts when an organism should cease foraging in one patch and travel to another.

mariculture The culture of marine organisms. In open mariculture (or semi-culture), organisms are cultured in natural environments; in closed mariculture (or intensive mariculture), organisms are cultured in a controlled environment.

marine Living in or pertaining to the sea.

marine archaeology The discovery, salvage, and interpretation of material remains of humankind's past that have been preserved in the sea.

marine natural products Chemical compounds that are obtained from marine organisms.

marine snow Detritus and other particulate organic matter that is found in the water column.

maritime culture A human culture with a very close relationship with the sea.

market equilibrium The dynamic balance between supply and demand under a given set of conditions in a "free" market (one with no monopolies or government interventions).

marrara Nasopharyngeal blockage by a parasite. Also called halzoun.

marsh Wetland without trees; in North America, this type of land is characterized by cattails and rushes.

marsupial One of the pouched mammals of the subclass Metatheria.

mass The fundamental unit of measurement equivalent to the weight of a substance when compared with the weight of hydrogen.

mass burn Incineration of unsorted solid waste.

masseter A chewer.

mass number Sum of the number of protons and neutrons in the nucleus of an atom; designated by a superscript to the upper left of the elemental symbol.

mast cell Type of cell in various tissues that releases pharmacologically active substances with a role in inflammation.

mastax The pharyngeal apparatus of rotifers used for grinding ingested food.

mastication Chewing.

mastigont Axoneme of a cilium or flagellum together with its basal fibrils and organelles.

mastigophora The protozoan subphylum where members possess one or more flagella that are used for locomotion; autotrophic, heterotrophic, or saprozoic.

mastitis Infection of the udder of cattle.

mastoid A bony process of the skull.

maternal Pertaining to the mother.

maternal condition model In polygynous mammals, females that are high ranking, well nourished, or otherwise in good condition giving birth to a greater proportion of sons since their sons will outreproduce their daughters and vice versa for females that are in poor condition.

mating system The species-typical pattern of mate finding, reproduction, and parenting of offspring.

matric potential The water potential component caused by the attraction of water molecules to a hydrophilic matrix.

matrix The intercellular substance of a tissue, or that part of a tissue into which an organ or process is set; site of the Krebs cycle.

matter Anything that has mass and occupies space.

maturation To complete the natural development of an animal system. Improvement in the performance of behavior occurs as parts of the nervous system and other structures complete development.

Maurer's clefts Blotches on the surface of an erythrocyte infected with Plasmodium falciparum.

maxilla One member of a pair of mouthparts located just posterior to the mandibles of many arthropods.

maxillae (second maxillae)Fifth pair of appendages in Crustacea, primarily feeding in function, derived from appendages on primitive sixth (third postoral) somite; homologous to labium in insects. The maxillae of insects are the third pair of head appendages, homologous to maxillules of Crustacea.

maxillary gland In malacostracan crustaceans, the excretory organs are located near the maxillary segments and are termed maxillary glands.

maxilliped One of the pairs of head appendages located just posterior to the maxilla in crustaceans, a thoracic appendage that has become incorporated into the feeding mouthparts.

maxillopodan eye Naupliar eye of crustacean class Maxillopoda; has a tapetum (crystalline reflective layer).

maxillules (first maxillae)Fourth pair of appendages in Crustacea, primarily feeding in function; derived from appendages on primitive fifth (second postoral) somite; homologous to maxillae in insects.

maximum sustainable yield The maximum level of fishing effort that a fish stock can withstand without causing major upsets in the abundance of its stock.

meadow An open clearing in a landscape in which low-growing, herbaceous plants dominate.

mean The central tendency of a series of measurements or observations, calculated by summing all values in a distribution and dividing the total by the number of values.

meatus A passageway.

mechanical vector Vector that transmits disease organism by mechanical means only. Contrast with biological vector.

mechanoreceptor A sensory receptor that is sensitive to mechanical stimulation, such as changes in pressure or tension.

meconium A dark green mucilaginous material in the intestine of the full-term fetus, being a mixture of the secretions of the intestinal glands and some amniotic fluid.

medial On or near the plane that divides a bilateral animal into mirror images. Also median.

median (parietal) eye A photoreceptor located middorsally on the head of some vertebrates; it is associated with the vertebrate epithalamus.

mediastinum The membranes enclosing the tissues and organs between the two lungs.

mediated transport. Transport of a substance across a cell membrane mediated by a carrier molecule in the membrane.

Mediterranean climate areas Specialized landscapes with warm, dry summers; cool, wet winters; many unique plant and animal adaptations; and many levels of endemism.

medulla oblongata Portion of the brain stem located between the pons and the spinal cord.

medulla The inner portion of an organ in contrast to the cortex or outer portion. Also, hindbrain.

medusa A jellyfish, or the free-swimming stage in the life cycle of cnidarians.

megacity See megalopolis.

megacolon Flabby distended colon caused by chronic Chagas' disease.

megaesophagus Distended esophagus caused by chronic Chagas' disease.

megagametogenesis The production of megagametes (large gametes) from megaspores in the ovules of angiosperms.

megagametophyte The gametophyte stage containing eight haploid nuclei within the embryo sac.

megalopolis Also known as a megacity or supercity; megalopolis indicates an urban area with more than 10 million inhabitants.

megaplankton The component of the plankton that consists of very large organisms over 20 cm in size.

megasporangium Sporangium that contains megaspores.

megaspore Haploid cell produced by meiosis in the ovules of angiosperms; a single megasporocyte produces four megaspores, only one of which remains functional.

megaspore mother cell Diploid cell in megasporangium that, upon undergoing meiosis, yields megaspores.

megasporocyte Also known as the megaspore mother cell; this diploid cell undergoes meiosis to produce four haploid megaspores in the ovules of angiosperms.

megasporogenesis The process of megaspores (large spores) being produced via meiosis in the ovules of angiosperms.

megawatt (MW) Unit of electrical power equal to one thousand kilowatts or one million watts.

Mehlis' glands Unicellular mucous and serous glands surrounding the ootype of a flatworm.

meiofauna Small invertebrates found in the interstices between sand grains.

meiosis Process of cell division by which egg and sperm cells are formed, involving a diminution in the amount of genetic material. Comprises two successive nuclear divisions with only one round of DNA replication, which produces four haploid daughter cells from an initial diploid cell.

meiotic Cell division that reduces the number of chromosomes from diploid to haploid.

Meissner'scorpuscles Tactile sensory receptors in the superficial dermis.

melanin Black or dark-brown pigment found in plant or animal structures.

melanism Dark coloration of the skin, fur, or feathers because of the presence of the pigment melanin.

melanophore Black or brown chromatophore containing melanin.

melatonin A hormone secreted by the pineal gland. Functions in regulating photoperiodicity.

melon A fatty structure on the forehead of some cetaceans that is used to direct sound waves emitted during echolocation.

membrane In living organisms, a phospholipid bilayer impregnated with protein and certain other compounds; functions in partitioning of cellulose activities.

membrane selectivity See differentially permeable membrane.

membranelle Short, transverse rows of cilia, fused at their bases, serving to move food particles toward the oral groove of a protozoan.

memory The capacity of an organism to form lasting connections based on past experiences; the ability to store and use information.

memory cell A lymphocyte capable of initiating the antibody- mediated immune response on detection of a specific antigen molecule for which it is genetically programmed. It circulates freely in the blood and lymph, and may live for years.

memory trace See engram.

meninges Any of three membranes (arachnoid, dura mater, pia mater) that envelop the vertebrate brain and spinal cord. Also, solid connective tissue sheath enclosing the central nervous system of some vertebrates.

menisci A crescent-shaped fibrocartilage.

menopause In the human female, that time of life when ovulation ceases; cessation of the menstrual cycle.

menses The period of shedding of the lining (endometrium) of the uterus and associated fluids if an ovum is not fertilized, most notably in primates.

menstrual cycle The period of the regularly recurring physiologic changes in the endometrium that culminates in its shedding (menstruation).

menstruation Loss of blood and tissue from the uterus at the end of a female primate's reproductive cycle.

meristem The tissue or zone from which new cells are produced by cell division.

meristematic tissue Within some seaweeds, specific tissue sites where most cell division for growth occurs.

meroblastic Partial cleavage occurring in zygotes having a large amount of yolk at the vegetal pole; cleavage restricted to a small area on the surface of the egg.

merogony Multiple fission to produce merozoites; schizogony.

meroplankton Planktonic organisms that spend only part of their life in the plankton. Compare holoplankton.

Merostomata The class of arthropods whose members are aquatic and possess book gills on the opisthosoma. Eurypterids (extinct) and horseshoe crabs.

merozoite A very small trophozoite at the stage just after cytokinesis has been completed in multiple fission of a protozoan.

mesenchyme Undifferentiated mesoderm. It will eventually develop into muscle, blood vessels, skeletal elements, and (other) connective tissues.

mesenterial filament Any of the long, thin tubes attached to the gut of corals and other cnidarians that are involved in digestion and absorption.

mesenteries Infoldings of gastroderm and mesoglea extending into the gastrovascular cavity of cnidarians; sheets of peritoneum from which the digestive tract is suspended in coelomates.

mesentery Peritoneal fold serving to hold the viscera in position.

mesic Referring to a region that receives adequate precipitation to maintain biological productivity.

mesocarp The middle layer of the fruit wall, located between the exocarp and endocarp.

mesocercaria Juvenile stage of the digenetic trematode Alaria. It is an unencysted form between the cercaria and the metacercaria.

mesocoel Middle body coelomic compartment in some deuterostomes, anterior in lophophorates, corresponds to hydrocoel in echinoderms.

mesoderm The third germ layer, formed in the gastrula between the ectoderm and endoderm; gives rise to connective tissues, muscle, urogenital and vascular systems, and the peritoneum.

mesoglea The layer of jellylike or cement material between the epidermis and gastrodermis in cnidarians and ctenophores; also may refer to jellylike matrix between epithelial layers in sponges.

mesohyl Gelatinous matrix surrounding sponge cells; mesoglea, mesenchyme.

mesolecithal Pertaining to a zygote (or ovum) having a moderate amount of yolk concentrated in the vegetal pole.

mesonephros The middle of three pairs of embryonic renal organs in vertebrates. Functional kidney of fishes and amphibians; its collecting duct is a Wolffian duct. Adj., mesonephric.

mesopelagic zone The pelagic environment from a depth of approximately 100 to 200 m (350 to 650 ft) to 1,000 m (3,000 ft).

mesophyll Photosynthetic middle layer in the blade of a leaf; typically composed of palisade and spongy parenchyma.

mesophyte A plant that grows in soils that contain moderate or intermediate amounts of moisture.

mesoplankton The component of the plankton that consists of organisms 0.2 to 2 mm in size .

mesosome The portion of the body in lophophorates and some deuterostomes that contains the mesocoel.

mesosphere The atmospheric layer above the stratosphere and below the thermosphere; the middle layer; temperatures are usually very low.

mesothorax The middle of the three thoracic segments of an insect; usually contains the second pair of legs and the first pair of wings.

Mesozoa A phylum of animals whose members are parasites of marine invertebrates. With a two-layered body organization. Dioecious, complex life histories. Orthonectids and dicyemids.

Mesozoic The geological era beginning about 250 million years ago. During this time gymnosperm forests, dinosaurs, mammals, birds, and flowering plants appeared. By the end of the Mesozoic, about 70 million years ago, dinosaurs became extinct.

messenger RNA A single-stranded polyribonucleotide; formed in the nucleus from a DNA template and carries the transcribed genetic code to the ribosome where the genetic code is translated into protein.

metabolic energy That energy obtained from ATP produced in metabolism.

metabolism A group of processes that includes digestion, production of energy (respiration), and synthesis of molecules and structures by organisms; the sum of the constructive (anabolic) and destructive (catabolic) processes.

metabolites Chemical substances required in metabolism.

metacentric Chromosome with centromere at or near the middle.

metacercaria Stage between the cercaria and adult in the life cycle of most digenetic trematodes; usually encysted and quiescent.

metacestode Developmental stage of a cestode after metamorphosis of the oncosphere; a juvenile cestode.

metacoel Posterior coelomic compartment in some deuterostomes and lophophorates; corresponds to somatocoel in echinoderms.

metacommunication Communication about communication, where one signal changes the meaning of signals that follow.

metacryptozoite Merozoite developed from a cryptozoite.

metacyclic Stage in the life cycle of a parasite that is infective to its definitive host.

metacyst Cystic stage of a parasite that is infective to a host.

metamere A repeated body unit along the longitudinal axis of an animal; a somite, or segment.

metamerism Division of the body along the anteroposterior axis into a serial succession of segments, each of which contains identical or similar representatives of all the organ systems of the body; primitively in arthropods, including externally a pair of appendages and internally a pair of nerve ganglia, a pair of nephridia, a pair of gonads, paired blood vessels and nerves, and a portion of the digestive and muscular systems.

metamorphic rock A type of rock, either granitic or sedimentary in origin, and structurally changed by high temperature and pressure.

metamorphosis Type of development in which one or more juvenile types differ markedly in body form from the adult; occurs in numerous animal phyla. Also applies to the actual process of changing from larval to adult form.

metanauplius Later naupliar larvae of some crustaceans; that is, occurring after several naupliar stages but before another larval type or preadult in the developmental sequence.

metanephridium An excretory organ found in many invertebrates; it consists of a tubule that has one end opening at the body wall and the opposite end in the form of a funnel-like structure that opens to the body cavity.

metanephros Embryonic renal organs of vertebrates arising behind the mesonephros; the functional kidney of reptiles, birds, and mammals. It is drained from a ureter.

metaphase Stage in mitosis when chromosomes become aligned in the middle of the cell and firmly attached to the mitotic spindle but have not yet segregated toward opposite poles.

metapodosoma Portion of the podosoma that bears the third and fourth pairs of legs of a tick or mite.

metapolar cells Posterior tier of cells in the calotte of a dicyemid mesozoan.

metasome The portion of the body in lophophorates and some deuterostomes that contains the metacoel.

metathorax The posterior of the three segments of an insect thorax; it usually contains the third pair of walking legs and the second pair of wings (Arthropoda).

metazoa Multicellular animals.

metazoan A multicellular animal.

methane CH4, natural gas.

methanogen A bacterium that obtains energy from CO2 and H2 and forms methane.

metraterm Muscular, distended termination of the uterus of a digenetic trematode.

MHC See major histocompatibility complex.

micelle Lipid aggregates with a surface coat of bile salts. A stage in the digestion of lipids in the small intestine.

micro-hydro generators Small power generators that can be used in low-level rivers to provide economical power for four to six homes, freeing them from dependence on large utilities and foreign energy supplies.

microatoll A small, flat, atoll-shaped coral structure generally found in protected coral lagoons.

microbial loop A component of epipelagic food webs in which dissolved organic matter is cycled through the picoplankton and nanoplankton back into the main part of the food web.

microbody Membrane-bound organelles that are the site of certain enzymatic conversions; example: peroxisomes and glyoxisomes.

microevolution A change in the gene pool of a population across generations.

microfauna benthic animals smaller than about 0.1 mm

microfibril An elongated strand of cellulose molecules.

microfilament Component of the cytoskeleton; involved in cell shape, motion, and growth. Helical protein filament formed by the polymerization of globular actin molecules.

microfilaria The prelarval stage of filarial worms. Found in the blood of humans and the tissues of the vector.

microfilariae Partially developed juveniles borne alive by filarial worms (phylum Nematoda).

microfossils The microscopic shells and other remains of marine organisms that make up biogenous sediments.

microgamete The smaller of the two gamete types in a heterogametic organism, considered the male gamete.

microgametocyte Cell that gives rise to microgametes.

microgametogenesis The production of microgametes (small gametes) from microspores in the anthers of flowering plants.

micromere The smallest size class of blastomeres in a cleaving embryo when the blastomeres differ in size from one another.

micron (m) One one-thousandth of a millimeter; about 1/25,000 of an inch. Now largely replaced by micrometer (mm).

microneme One of the types of structures composing the apical complex in the phylum Apicomplexa, slender and elongate, leading to the anterior and thought to function in host cell penetration.

micronemes Slender, convoluted bodies that join a duct system with the rhoptries, opening at the tip of a sporozoite or merozoite.

microniscus Intermediate larval stages of the isopod suborder Epicaridea, parasitic on free-living copepods.

micronucleus A small body of DNA that contains the hereditary information of ciliates (Protista); exchanged between protists during conjugation. It undergoes meiosis before functioning in sexual reproduction.

micronutrient A dietary element essential in only small quantities (e.g., iron, chlorine, copper, and vitamins).

microorganisms Single-celled living organisms that can be seen only with the aid of a microscope.

microparasite Small (or very small) parasite that multiplies within the host of interest. Examples are protistan and prokaryotic parasites.

microplankton The component of the plankton that consists of organisms 20 to 200 mm (0.02 to 0.2 mm) in size.

micropores Spaces between soil particles that hold water by means of capillary forces.

micropredator Temporary parasite.

micropyle The small opening through which the cells emerge from a gemmule (phylum Porifera).

microscopy Examination with a microscope.

microspora The protozoan phylum characterized by members having unicellular spores; intracellular parasites in nearly all major animal groups. Examples: microsporeans (Nosema).

microsporangium The structure (sporangium) in which microspores are produced.

microspore Haploid cells produced by meiosis in the anthers of angiosperms; four microspores are produced from a single microsporocyte.

microspore mother cell Diploid cell in microsporangium that undergoes meiosis to produce microspores.

microsporocyte Also known as the microspore mother cell, this diploid cell undergoes meiosis to produce four haploid microspores in the anthers of angiosperms.

microsporogenesis The process of microspores (small spores) being produced via meiosis in the anthers of angiosperms.

microsporophyll A leaflike structure giving rise to one or more microsporangia.

microthrix (microtriches) Minute projections of the tegument of a cestode.

microtubule A hollow cylinder of tubulin subunits; involved in cell shape, motion, and growth; functional unit of cilia and flagella. It is one of three major classes of filaments of the cytoskeleton.

microvillus Narrow, cylindrical cytoplasmic projection from epithelial cells; microvilli form the brush border of several types of epithelial cells. Also, microvilli with unusual structure cover the surface of cestode tegument (also called microthrix [pl. microtriches]).

mictic Pertaining to haploid egg of rotifers or the females that lay such eggs.

mictic eggs Pertaining to the haploid eggs of rotifers. If it isn't fertilized, the egg develops parthenogenetically into a male; if fertilized, mictic eggs secrete a heavy shell and become dormant, hatching in the spring into amictic females.

midbrain The portion of the brain between the pons and forebrain.

mid-ocean ridge The continuous chain of volcanic submarine mountains that extends around the earth. It includes the Mid-Atlantic Ridge and East Pacific Rise.

middle lamella Layer of adhesive material (primarily pectins) found between adjacent cell walls.

midrib The central large vein of a leaf.

midwater Pertaining to the mesopelagic zone.

Miescher's tubules Sarcocysts; tissue cysts of Sarcocystis.

migration A seasonal movement from one location to another.

Milankovitch cycles Periodic variations in tilt, eccentricity, and wobble in the earth's orbit; Milutin Milankovitch suggested that it is responsible for cyclic weather changes.

milpa agriculture An ancient farming system in which small patches of tropical forests are cleared and perennial polyculture agriculture practiced and is then followed by many years of fallow to restore the soil; also called swidden agriculture.

mimicry When one species resembles one or more other species; often protection is afforded the mimic species.

mineral A naturally occurring, inorganic, crystalline solid with definite chemical composition and characteristic physical properties.

mineralocorticoids Hormones of the adrenal cortex, especially aldosterone, that regulate salt balance.

miracidium The ciliated, free-swimming first stage larva of a digenean trematode that undergoes further development in the body of a snail.

Mississippian Part of the Carboniferous period, beginning around 360 million years ago.

mitigation Repairing or rehabilitating a damaged ecosystem or compensating for damage by providing a substitute or replacement area.

mitochondria in eucaryotes, a subcellular organelle that conducts cellular respiration

mitochondrion Membrane-bounded organelle that specializes in aerobic respiration (oxidative phosphorylation) and produces most of the ATP in eukaryotic cells.

mitosis Nuclear division in which there is an equal qualitative and quantitative division of the chromosomal material between the two resulting nuclei; ordinary cell division.

mitotic apparatus Collectively, the asters, spindle, centrioles, and microtubules of a dividing cell.

mixed perennial polyculture Growing a mixture of different perennial crop species (where the same plant persists for more than one year) together in the same plot; imitates the diversity of a natural system and is often more stable and more suitable for sustainable agriculture than monoculture of annual plants.

mixed semidiurnal tide A tidal pattern with two successive high tides of different heights each day.

modal action pattern (MAP) A spatiotemporal behavior pattern that is common to members of a species; different individuals perform the pattern in a recognizably similar fashion.

modern synthesis The combination of principles of population genetics and Darwinian evolutionary theory.

modification A change, as in morphology, usually associated with a functional advantage.

molar L. molaris = a grinder.

mole One gram molecular weight of any substance; that is, the molecular weight of any substance in grams.

molecule A particle composed of two or more atoms bonded together. An aggregate of at least two atoms in a definite arrangement held together by special forces.

molecular biology The study of the biochemical structure and function of organisms.

molecular genetics The study of the biochemical structure and function of DNA.

molecular weight The sum of all the atomic weights of a molecule.

Mollusca The phylum of coelomate animals whose members possess a head-foot, visceral mass, mantle, and mantle cavity. Most molluscs also possess a radula and a shell. The molluscs. Bivalves, snails, octopuses, and related animals.

molluscs Invertebrates with a soft, unsegmented body, a muscular foot, and, with some exceptions, a calcareous shell.

molt A process in arthropods, such as crustaceans and insects, as well as in some vertebrates, such as snakes, whereby the organism sheds its outer exoskeleton or skin periodically as it grows. Also refers to changing the pelage or feathers in a mammal or bird.

molt The exoskeleton that is shed during the molting process.

molting See ecdysis.

Monera The kingdom of life whose members are characterized by having cells that lack a membrane-bound nucleus, as well as other internal, membrane-bound organelles (they are prokaryotic); bacteria.

monerans Members of the kingdom Monera, which consists of prokaryotic organisms.

monestrous A species in which the female is receptive for only a few days once each year.

monitored, retrievable storage Holding wastes in underground mines or secure surface facilities such as dry casks where they can be watched and repackaged, if necessary.

monkey wrenching Environmental sabotage such as driving large spikes in trees to protect them from loggers, vandalizing construction equipment, pulling up survey stakes for unwanted developments, and destroying billboards.

monocarpic Denoting a plant that flowers only once.

monocot One of the two primary groups of angiosperms characterized by a single cotyledon, parallel venation of leaves, and floral parts in threes.

monocotyledon A class of angiosperms in which the seedlings typically possess one cotyledon. Commonly abbreviated to monocot.

monoculture An agricultural system in which only one crop species is cultivated.

monoculture agroforestry Intensive planting of a single species; an efficient wood production approach, but one that encourages pests and disease infestations and conflicts with wildlife habitat or recreation uses.

monocyte A type of leukocyte that becomes a phagocytic cell (macrophage) after moving into tissues.

monoecious Hermaphroditic; an individual that contains reproductive systems of both sexes.

monogamous Having one mate at a time.

monogamy The condition of having a single mate at any one time.

Monogenea The class of Platyhelminthes that has members that are called monogenetic flukes; most ectoparasites on vertebrates (usually on fishes, occasionally on turtles, frogs, copepods, squids); one life-cycle form in only one host; bear an opisthaptor. Examples: Disocotyle, Gyrodactylus, Polystoma.

Monogononta A class of rotifers containing members that possess one ovary; mastax not designed for grinding; produce mictic and amictic eggs. Example: Notommata.

monohybrid cross A mating between two individuals heterozygous for one particular trait.

monohybrid A hybrid offspring of parents different in one specified character.

monomer A molecule of simple structure, but capable of linking with others to form polymers.

monophyletic groups Groups that contain an ancestor and all the descendants of that ancestor.

monophyletic origin A group of organisms that evolved from a single ancestral type.

monophyly The condition that a taxon or other group of organisms contains the most recent common ancestor of the group and all of its descendants; contrasts with polyphyly and paraphyly.

Monoplacophora The class of molluscs whose members have a single, arched shell; a broad, flat foot; and certain serially repeated structures. Neopilina.

monoplacophorans A small group of molluscs that are thought by some to represent a link with invertebrates that show segmentation.

monosaccharide A simple sugar that cannot be decomposed into smaller sugar molecules; the most common are pentoses (such as ribose) and hexoses (such as glucose).

monostome Fluke that lacks a ventral sucker.

monounsaturated fat Composed of fatty acid chains in which there is only a single C-C double bond; examples: canola oil and olive oil.

monoxenous Living within a single host during a parasite's life cycle.

monozoic Tapeworms with a single proglottid, do not undergo strobilation to form chain of proglottids.

monozygotic Twins that arise from a single zygote, hence, two genetically identical individuals.

monsoon A seasonal reversal of wind patterns caused by the different heating and cooling rates of the oceans and continents.

montane coniferous forests Coniferous forests of the mountains consisting of belts of different forest communities along an altitudinal gradient.

moral agents Beings capable of making distinctions between right and wrong and acting accordingly. Those whom we hold responsible for their actions.

moral subjects Beings that are not capable of distinguishing between right or wrong or that are not able to act on moral principles and yet are susceptible of being wronged by others. This category assumes some rights or inherent values in moral subjects that gives us duties or obligations towards them.

morbidity Illness or disease.

more-developed countries (MDC) Industrialized nations characterized by high per capita incomes, low birth and death rates, low population growth rates, and high levels of industrialization and urbanization.

morphine A pain-relieving and addictive compound derived from the opium poppy (Papaver somniferum).

morphogenesis Development of the architectural features of organisms; formation and differentiation of tissues and organs.

morphology The science of structure. Includes cytology, the study of cell structure; histology, the study of tissue structure; and anatomy, the study of gross structure.

mortality Death rate in a population; the probability of dying.

morula A stage in the embryonic development of some animals that consists of a solid ball of cells.

mosaic An organism whose tissues are made up of two or more genetically different types.

mosaic cleavage. Embryonic development characterized by independent differentiation of each part of the embryo; determinate cleavage.

mosaic evolution A change in a portion of an organism (e.g., a bird wing) while the basic form of the organism is retained.

moss A group of terrestrial, nonvascular plants; the dominant plant body is the gametophyte with the sporophyte embedded in it.

motility Ability to move.

motivation Internal processes that arouse and direct behavior. Today this refers more to examination of the consequences of behavior and feedback to the animal concerning the consequences of those actions.

motor (efferent) neuron or nerve A neuron or nerve that transmits impulses from the central nervous system to an effector such as a muscle or gland.

motor neuron A neuron that synapses with a muscle membrane.

motor unit A motor neuron and the muscle fibers associated with it.

mountain tundra That portion of tundra vegetation confined to alpine meadows. Low-growing grasses, sedges, and forbs with a very short growing season; permafrost is typical.

mucigel A slimy material secreted by and covering the root cap and root hairs.

mucilaginous Containing a mucilage, usually composed of mucopolysaccharides.

mucin Any of a group of glycoproteins secreted by certain cells, especially those of salivary glands.

mucous cell A glandular cell that secretes mucus.

mucron Apical anchoring device on an acephaline gregarine protozoan.

mucus Viscid, slippery secretion rich in mucins produced by secretory cells such as those in mucous membranes. Adj., mucous.

mud flat A muddy bottom that is exposed at low tide.

Muellerian mimicry Evolution by one species to resemble the coloration, body shape, or behavior of an unrelated species that is protected from predators by a venomous stinger, bad taste, or some other adaptation.

mulch Protective ground cover, including manure, wood chips, straw, seaweed, leaves, and other natural products, or synthetic materials, such as heavy paper or plastic, that protect the soil, save water, and prevent weed growth.

Mullerian mimicry Occurs when two similar species are both distasteful to predators.

Mullerian mimics Noxious species that resemble each other.

Muller's larva A free-swimming ciliated larva that resembles a modified ctenophore, characteristic of many marine polyclad turbellarians.

Muller's ratchet The steady accumulation of mutations in a population of asexual organisms over time.

Müller's larva. Free-swimming ciliated larva that resembles a modified ctenophore, characteristic of certain marine polyclad turbellarians.

multigenic (polygenic) inheritance Inheritance in which the genetic control for a trait results in the phenotypic expression varying continuously.

multiple alleles A condition in which more than two alleles exist for a given trait.

multiple fission Asexual reproduction by the splitting of a cell or organism into many cells or organisms. See schizogony.

multiple fruit A fruit derived from the fusion of the ovaries of several flowers in an inflorescence; example: pineapple.

multiple use Many uses that occur simultaneously; used in forest management; limited to mutually compatible uses.

muscle fiber The contractile unit of a muscle.

muscle tissue The type of tissue that allows movement. The three kinds are skeletal, smooth, and cardiac. Tissue made of bundles of long cells called muscle fibers.

muscleL. musculus = a contractile organ.

muscularis mucosae Smooth muscle fibers around the mucosa of the gut wall, surrounding the lamina propria and surrounded by the submucosa.

mushroom Common name for a group of fungi (Basidiomycetes) that produce an aboveground reproductive structure.

musk gland See scent gland.

mutagen Any substance capable of causing a mutation.

mutagens Agents, such as chemicals or radiation, that damage or alter genetic material (DNA) in cells.

mutation pressure The constant resupplying of mutations to a gene pool due to a base mutation rate.

mutation A change, either spontaneous or by external factors, in the genetic material of a cell; mutations in the gametes (sex cells) can be inherited by future generations of organisms.

mutational analysis An analysis that examines the effects of specific mutants for the effects on particular phenomena with respect to morphology, physiology, or behavior.

mutualism A type of interaction in which two different species derive benefit from their association and in which the association is necessary to both; often symbiotic.

mycelium A network of fungal hyphae.

mycetome Specialized organ in some insects that bears mutualistic bacteria.

mycobiont The fungal partner in a mutualistic relationship such as mycorrhizae or lichens.

mycologist A person who studies fungi.

mycorrhiza Symbiotic association between a fungus and a plant root.

mycotoxin A toxic compound formed by the hyphae of common molds growing under a variety of conditions, especially in contaminated foods.

myelin A fatty material forming the medullary sheath of nerve fibers.

myiasis Infection by fly maggots.

myocyte Contractile cell (pinacocyte) in sponges.

myofibril A contractile filament within muscle or muscle fiber.

myogenic Originating in muscle, such as heartbeat arising in vertebrate cardiac muscle because of inherent rhythmical properties of muscle rather than because of neural stimuli.

myoglobin The oxygen-transporting pigment of muscle tissue.

myomere The muscle plate or portion of a somite that develops into voluntary muscle.

myosin A large protein of contractile tissue that forms the thick myofilaments of striated muscle. During contraction it combines with actin to form actomyosin.

myotome That part of a somite destined to form muscles; the muscle group innervated by a single spinal nerve.

myriapods Members of the four noninsect classes of the subphylum Uniramia. Includes centipedes, millipedes, pauropods, and symphylans.

Myxini The class of vertebrates whose members are fishlike, jawless, without paired appendages, and possess four pairs of tentacles around the mouth. Hagfishes.

Myxozoa The protozoan phylum characterized by members having spores of multicellular origin; the myxozoans.

myzorhynchus Apical stalked, suckerlike organ on the scolex of some tetraphyllidean cestodes.


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